Apis mellifera ligustica 4 Honeybee 4 honeybee 4 scent marking 4 Animalia 2 Apis mellifera 2 Apoidea 2 Colour vision 2 Communication 2 Efficiency 2 Foraging 2 Foraging behaviour 2 Foraging efficiency 2 Honeybees 2 Movement pattern 2 Private information 2 Scent marking 2 Scent-mark 2 Social information 2 Waggle dance 2 animal communication 2 animal experiment 2 article 2 bee 2 communication 2 controlled study 2 flower 2 food 2 forager 2 foraging 2