Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae

Hegetotheriidae is one of the most derived families of South American notoungulates, which are known from the late Oligocene to Pliocene. Several genera of hegetotheriids have been described; most of them have been recently re-examined, but the Miocene taxa Hegetotherium and Pachyrukhos have not bee...

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Detalles Bibliográficos
Publicado: 2019
Materias:
Argentina
Hegetotheriidae
Miocene
Notoungulata
systematics
Typotheria
Acceso en línea:https://bibliotecadigital.exactas.uba.ar/collection/paper/document/paper_14772019_v_n_p_Seoane
http://hdl.handle.net/20.500.12110/paper_14772019_v_n_p_Seoane
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Biblioteca Digital - Facultad de Ciencias Exactas y Naturales (UBA) de Universidad de Buenos Aires
id paper:paper_14772019_v_n_p_Seoane
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spelling paper:paper_14772019_v_n_p_Seoane2023-06-08T16:18:10Z Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae Argentina Hegetotheriidae Miocene Notoungulata systematics Typotheria Hegetotheriidae is one of the most derived families of South American notoungulates, which are known from the late Oligocene to Pliocene. Several genera of hegetotheriids have been described; most of them have been recently re-examined, but the Miocene taxa Hegetotherium and Pachyrukhos have not been revised for more than a century. Numerous species of both genera were described mainly from the Santa Cruz Formation, Santa Cruz Province, but also from the Colhuehuapian beds in Chubut Province, and the Colloncuran beds in Neuquén and Río Negro provinces, Argentina. Recently, Hegetotherium was recorded in Bolivia and Chile. Sinclair was the first reviewer to postulate some synonymies, in the first decade of the twentieth century. The present contribution provides a re-evaluation of these genera, including the study of previously unpublished materials. We conclude that Hegetotherium mirabile and Pachyrukhos moyani are the only hegetotheres in the Santa Cruz Formation (Santacrucian SALMA) with Hegetotherium novum and Pachyrukhos politus in the Colhuehuapian SALMA. Additionally, H. convexum, H. anceps, H. minum and H. andinum are proposed as synonyms of H. mirabile; H. novum is removed from Prohegetotherium; and Hegetotherium arctum is excluded from Hegetotheriidae. This taxonomic revision led to a new phylogenetic analysis of Hegetotheriidae, based on a previous data matrix, but enlarged and corrected, as some taxonomic issues concerning other genera were revealed. The analysis supports the monophyly of Hegetotheriidae and the subfamily Pachyrukhinae, as well as the monophyly of the genera Hemihegetotherium and Paedotherium, if the proposed removal of Pd. kakai from the latter is admitted. In contrast, Hegetotheriinae and Hegetotherium appear to be paraphyletic. The position of H. novum supports its removal from the genus Prohegetotherium, in which it was recently included. © 2019, © The Trustees of the Natural History Museum, London 2019. All rights reserved. 2019 https://bibliotecadigital.exactas.uba.ar/collection/paper/document/paper_14772019_v_n_p_Seoane http://hdl.handle.net/20.500.12110/paper_14772019_v_n_p_Seoane
institution Universidad de Buenos Aires
institution_str I-28
repository_str R-134
collection Biblioteca Digital - Facultad de Ciencias Exactas y Naturales (UBA)
topic Argentina
Hegetotheriidae
Miocene
Notoungulata
systematics
Typotheria
spellingShingle Argentina
Hegetotheriidae
Miocene
Notoungulata
systematics
Typotheria
Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae
topic_facet Argentina
Hegetotheriidae
Miocene
Notoungulata
systematics
Typotheria
description Hegetotheriidae is one of the most derived families of South American notoungulates, which are known from the late Oligocene to Pliocene. Several genera of hegetotheriids have been described; most of them have been recently re-examined, but the Miocene taxa Hegetotherium and Pachyrukhos have not been revised for more than a century. Numerous species of both genera were described mainly from the Santa Cruz Formation, Santa Cruz Province, but also from the Colhuehuapian beds in Chubut Province, and the Colloncuran beds in Neuquén and Río Negro provinces, Argentina. Recently, Hegetotherium was recorded in Bolivia and Chile. Sinclair was the first reviewer to postulate some synonymies, in the first decade of the twentieth century. The present contribution provides a re-evaluation of these genera, including the study of previously unpublished materials. We conclude that Hegetotherium mirabile and Pachyrukhos moyani are the only hegetotheres in the Santa Cruz Formation (Santacrucian SALMA) with Hegetotherium novum and Pachyrukhos politus in the Colhuehuapian SALMA. Additionally, H. convexum, H. anceps, H. minum and H. andinum are proposed as synonyms of H. mirabile; H. novum is removed from Prohegetotherium; and Hegetotherium arctum is excluded from Hegetotheriidae. This taxonomic revision led to a new phylogenetic analysis of Hegetotheriidae, based on a previous data matrix, but enlarged and corrected, as some taxonomic issues concerning other genera were revealed. The analysis supports the monophyly of Hegetotheriidae and the subfamily Pachyrukhinae, as well as the monophyly of the genera Hemihegetotherium and Paedotherium, if the proposed removal of Pd. kakai from the latter is admitted. In contrast, Hegetotheriinae and Hegetotherium appear to be paraphyletic. The position of H. novum supports its removal from the genus Prohegetotherium, in which it was recently included. © 2019, © The Trustees of the Natural History Museum, London 2019. All rights reserved.
title Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae
title_short Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae
title_full Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae
title_fullStr Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae
title_full_unstemmed Systematic revision of Hegetotherium and Pachyrukhos (Hegetotheriidae, Notoungulata) and a new phylogenetic analysis of Hegetotheriidae
title_sort systematic revision of hegetotherium and pachyrukhos (hegetotheriidae, notoungulata) and a new phylogenetic analysis of hegetotheriidae
publishDate 2019
url https://bibliotecadigital.exactas.uba.ar/collection/paper/document/paper_14772019_v_n_p_Seoane
http://hdl.handle.net/20.500.12110/paper_14772019_v_n_p_Seoane
_version_ 1768545480520761344

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